When former Princeton music professor Arthur Cochran unexpectedly stumbles into choir practice at a small-town church, he finds a group of singers that are out of tune in more ways than one. Despite their differences, he and his new cohorts may just help each other reinvent.
Arthur defies local customs, and the choir pays for his rudeness. For Cash's sake, Ginny vows to say nothing bad about Wayne.
Arthur helps Ginny conquer stage fright. Reverend Jax is caught in the rift between Wayne and Dwayne.
When Arthur is faced with a choir rebellion, he decides – against Ginny's advice – to take on Adams, the group's alpha, at their annual deer hunting trip. Wayne takes Cash on his first hunting trip to teach him how to be a man.
Arthur deals with Jean's father, who attempts to take their family houseboat back. Reverend Jax hosts a singles night, where Ginny and Dwayne finally address the elephant in their relationship.
When Ginny tries to step into a new management role at work, Arthur helps her overcome her fears of inadequacy. When Dwayne opts out of a longstanding Halloween tradition, Wayne fears that they are growing apart.
After Pastor Magnus invites the Second First Choir to perform at the Church of Perpetual Praise, Arthur resolves to sound better than ever to spite him. Ginny spirals when Arthur adds a new singer from her past to the choir.
Reverend Jax asks Arthur to watch the church for him. As parishioners come to him with problems, Arthur believes he can best help them with reason rather than with the ways of the church. Jax also asks Wayne to pick up Adams from the hospital.
Jax convinces Arthur to join the choir's progressive Thanksgiving dinner and snoop in choir members' homes to find a missing item. Meanwhile, Adams invites her handsome contractor to dinner as a set-up for Ginny, but he has something else in mind.